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Nelson, S.M., M. L. Niemiller, & B.M. Fitzpatrick. 2017. Co-occurrence and hybridization between Necturus maculosus and a heretofore unknown Necturus in the Southern Appalachians. Journal of Herpetology 51: 559-566.

Dresser, C.M.*, R.M. Ogle, & B.M. Fitzpatrick. 2017. Genome scale assessment of a species translocation program. Conservation Genetics doi:10.1007/s10592-017-0970-6

Marion, Z.H.*, J.A. Fordyce, & B.M. Fitzpatrick. 2017. Pairwise beta diversity resolves an underappreciated source of confusion in diversity partitioning. Ecology 98: 933-939.

Pierson,  TW*,  SJ  Arnold,  MK  Hamed,  W  Lattea,  & ET  Carter*(in  press)  Courtship behavior  of  the  Yonahlossee  Salamander  (Plethodon  yonahlossee): Observations  in the field and laboratory. Herpetological Conservation and Biology

Literman, R., S. Radhakrishnan, J. Tamplin, R. Burke, C. Dresser*, and N. Valenzuela (2017) Development of sexing markers in Glyptemys insculpta and Apalone spinifera turtles uncovers an XX/XY sex-determining system in the critically-endangered bog turtle Glyptemys muhlenbergii. Conservation Genetics Resources.DOI: 10.1007/s12686-017-0711-7.


Carter, ET*, MJ Ravesi, BC Eads, & BA Kingsbury. 2016. Invasive plant management creates ecological traps for snakes.Biological Invasions19(1): 443-453.

Attum, O, J Lowry, BA Kingsbury, &ET Carter*. 2016. Thedistribution of amur honeysuckle and box turtle habitat use in an urban forest.Urban Naturalist12: 1-16.

Niemiller, ML, KS Zigler, DR Stephen, ET Carter, A Paterson, SL Taylor, & AS Engel (2016) Vertebrate fauna in caves of the eastern Tennessee Appalachian Valley and Ridge region, USA. Journal of Cave and Karst Studies 78(1): 1-24.


Marion ZH, Fordyce JA, Fitzpatrick BM. 2015. Extending the concept of diversity partitioning to characterize phenotypic complexity. American Naturalist 186: 348-361.

Fitzpatrick BM, Ryan ME, Johnson JR, Corush J, Carter ET. 2015. Hybridization and the species problem in conservation. Current Zoology 61: 206-216.

Carter, ET, BC Eads, & BA Kingsbury (2015) Exotic Invasive Plants Alter Thermal Regimes: Implications for Management Using a Case study of a Native Ectotherm. Functional Ecology 29(5): 683-693.

Carter, ET, O Attum, BC Eads, AS Hoffman, & BA Kingsbury (2014) Reducing the Potential for Human-Snake Encounters in a Recreational Park. Human-Wildlife Interactions 8(2): 158-167. (cover article)

Jordan, MA, N Perrine-Ripplinger , & ET Carter (2014) An Independent Observation of Facultative Parthenogenesis in the Copperhead (Agkistrodon contortrix). Journal of Herpetology 49(1): 118-121.

Owen, Dustin, ET Carter, ML Holding, IT Moore, & K Islam (2014) Roads are Associated with a Blunted Stress Response in a North American Pit-Viper. General and Comparative Endocrinology 202(1): 87-92.


FItzpatrick, B. M. and A. L. Allison. 2014. Similarity and differentiation between bacteria associated with skin of salamanders (Plethodon jordani) and free-living assemblages. FEMS Microbiology Ecology DOI: 10.1111/1574-6941.12314

Fordyce, JA, Shah P, Fitzpatrick BM. 2014. iteRates: An R Package for Implementing a Parametric Rate Comparison on Phylogenetic Trees. Evolutionary Bioinformatics 2014:10 127-130 doi: 10.4137/EBO.S16487

Fitzpatrick, B. M. 2014. Symbiote transmission and maintenance of extra-genomic associations. Frontiers in Microbiology 5. doi: 10.3389/fmicb.2014.00046


Fitzpatrick, B. M. 2013. Alternative forms for genomic clines. Ecology and Evolution  3: 1951-1966

Dittrich-Reed, D. R. and B. M. Fitzpatrick. 2013. Transgressive hybrids as hopeful monsters. Evolutionary Biology 40: 310-315.

Niemiller, M. L., McCandless, J. R., Reynolds, R. G., Caddle, J., Near, T. J., Tillquist, C. R., Pearson, W. D., and Fitzpatrick, B. M. 2013. Effects of climatic and geological processes during the pleistocene on the evolutionary history of the northern cavefish, amblyopsis spelaea (teleostei: Amblyopsidae). Evolution 67: 1011-1025.

Reynolds, R. G. and B. M. Fitzpatrick. Tests of two methods for identifying founder effects in metapopulations reveal substantial type II error. Genetica 141: 119-131.

Shah, P., B. M. Fitzpatrick, J. A. Fordyce. 2013. A parametric method for assessing diversification rate variation in phylogenetic trees. Evolution 67: 368-377.

Ryan, M. E., J. R. Johnson, L. Lowenstine, A. Picco, B. M. Fitzpatrick, H. B. Shaffer. 2013. Lethal effects of water quality on threatened California salamanders but not on co-occurring hybrid salamanders. Conservation Biology 27: 95-102.

Niemiller ML, Fitzpatrick BM, Shah P, Schmitz L, and Near TJ. 2013. Evidence for repeated loss of selective constraint in rhodopsin of amblyopsid cavefishes (Teleostei: Amblyopsidae). Evolution 67: 732-748.

Barun A, Niemiller ML, Fitzpatrick BM, Fordyce JA, and Simberloff D. Can genetic data confirm or refute historical records: the island invasion of the small Indian mongoose (Herpestes auropunctatus). Biological Invasions. In Press


Reynolds, R. G., W. Booth, G. W. Schuett, B. M. Fitzpatrick, and G. M. Burghardt. 2012. Successive virgin births of viable male progeny in the checkered gartersnake, Thamnophis marcianus. Biological Journal of the Linnean Society 107:566-572.

Fitzpatrick, B. M. 2012. Estimating ancestry and heterozygosity of hybrids using molecular markers. BMC Evolutionary Biology 12: 131.

Fitzpatrick, B. M. 2012. Underappreciated consequences of phenotypic plasticity for ecological speciation. International Journal of Ecology (special issue on ecological speciation) 2012:256017 doi:10.1155/2012/256017

Reynolds, R. G., M. L. Niemiller, and B. M. Fitzpatrick. 2012. Genetic analysis of an endemic archipelagic lizard reveals sympatric cryptic lineages and taxonomic discordance. Conservation Genetics 13:953-963.

Placyk, J. S., B. M. Fitzpatrick, G. S. Casper, R. L. Small, R. G. Reynolds, D. W. A. Noble, R. J. Brooks, and G. M. Burghardt. 2012. Hybridization between two gartersnake species (Thamnophis) of conservation concern: a threat or an important natural interaction? Conservation Genetics 13:649-663.

Fitzpatrick, B.M., J.A. Fordyce, M. Neimiller, & R.G. Reynolds. 2012. What can DNA tell us about biological invasions? Biological Invasions. 14:245-253 PDF

Niemiller, M. L., T. J. Near and B. M. Fitzpatrick. 2012. Delimiting species using multilocus data: diagnosing cryptic diversity in the Southern Cavefish, Typhlichthys subterraneus (Teleostei: Amblyopsidae). Evolution. 66:846-866. DOI: 10.1111/j.1558-5646.2011.01480.x


Reynolds, RG, GP Gerber, and BM Fitzpatrick. 2011. Unexpected shallow genetic divergence in Turks Island Boas (Epicrates c. chrysogaster) reveals single evolutionary significant unit for conservation. Herpetologica. 67: 477-486.

Pasachnik SA, Echternacht AC, Fitzpatrick BM. 2011. Population genetics of the Honduran spiny-tailed iguana Ctenosaura melanosterna: implications for conservation and management. Endangered Species Research 14:113-126.

Bailey, J.K., Schweitzer, J.A., Úbeda, F., Zinkgraf, M., Fitzpatrick, B.M., O’Reilly-Wapstra, J., Rehill, B.J., LeRoy, C.J., Potts, B.M., Whitham, T.G., Genung, M.A., Fischer, D.G., Pregitzer, C.C., & Keith, A. 2011. From Genes to Ecosystems: Emerging Concepts Bridging Ecological and Evolutionary Dynamics. Ecological Reviews. The ecology of plant secondary metabolites: from genes to landscapes. Lason GR, Dicke M, and Hartley SE. Cambridge Univ Press, Cambridge, UK.

Genung, M. A., F. Ubeda, B. M. Fitzpatrick, J. A. Schweitzer, C. C. Pregitzer, E. Felker-Quinn, and J. K. Bailey. 2011. Genetic variation and community change – selection, evolution, and feedbacks. Functional Ecology 25:408-419. PDF


Reynolds, R.G. In Press. Status, conservation, and introduction of amphibians and reptiles in the Turks and Caicos            Islands, British West Indies. In: A. Hailey, B.S. Wilson, and J.A. Horrocks (eds.). Conservation of Caribbean Island            Herpetofaunas. Vol. II. Brill, Netherlands.

Reynolds, R.G., M.L. Niemiller, and S.A. Pasachnik. In Press. Pygmy Salamanders (Desmognathus wrighti) in            Tennessee. Journal of the Tennessee Academy of Science.

Niemiller, M.L. and R.G. Reynolds (eds). In Press. The Amphibians of Tennessee. The University of Tennessee Press,            Knoxville,Tennessee.

Niemiller, M.L.,and V.A. Cobb. In Press. Feeding activity of Amphiuma tridactylum at Reelfoot Lake, Tennessee.          Journal of the Tennessee Academy of Science.

Hadziabdic, D., B.M. Fitzpatrick, X. Wang, P.A. Wadl, T.A. Rinehart, B.H. Ownley, M.T. Windham, and R.N. Trigiano.            2010. Analysis of genetic diversity in flowering dogwood natural stands using microsatellites: the effects of            dogwood anthracnose. Genetica 138: 1047-1057.

Johnson, J.R., B.M. Fitzpatrick, and H.B. Shaffer. 2010. Retention of low-fitness genotypes over six decades of            admixture between native and introduced tiger salamanders. BMC Evolutionary Biology 10:147.

Reynolds, R.G. and M.L. Niemiller. 2010. Island invaders: introduced amphibians and reptiles in the Turks and Caicos          Islands. Reptiles and Amphibians 17: 117-121.   PDF

Pasachnik, S.A., B.M. Fitzpatrick, and A.C. Echternacht. 2010. Gene trees, species, and species trees in the          Ctenosaura palearis clade. Conservation Genetics 11: 1767-1781.

Niemiller, M.L.,and B.T. Miller. 2010. Gyrinophilus gulolineatus. Catalogue of American Amphibians and Reptiles.
         862.1 PDF

Fitzpatrick, B.M., J.R. Johnson, D.K. Kump, J.J. Smith, S.R. Voss, and H.B. Shaffer. 2010. Rapid spread of invasive genes          into a threatened native species. Proceedings of the National Academy of Sciences, USA. 107:3606-3610.          PDF

Mims, M.C., C.D. Hulsey, B.M. Fitzpatrick, and J.T. Streelman. 2010. Geography disentangles introgression from          ancestral polymorphism in Lake Malawi cichlids. Molecular Ecology 19: 940-951.   PDF

Niemiller, M.L., and T.L. Poulson. 2010. Studies of the Amblyopsidae: past, present, and future. In: E. Trajano, M.E.          Bichuette & B.G. Kapoor (eds.): The Biology of Subterranean Fishes. Science Publishers, Enfield, NH, USA.

Niemiller, M.L., M.S. Osbourn, D.B. Fenolio, T.K. Pauley, B.T. Miller, J.R. Holsinger. 2010. Conservation status and          habitat use of the West Virginia Spring Salamander (Gyrinophilus subterraneus) and Spring Salamander (G.          porphyriticus) in General Davis Cave, Greenbrier Co., West Virginia. Herpetological Conservation and Biology.
         5: 32-43.   PDF

Graening, G.O., D.B. Fenolio, M.L. Niemiller, A.V. Brown, and J.B. Beard. 2010. The 30-year recovery effort for the Ozark          cavefish (Amblyopsis rosae): analysis of current distribution, population trends, and conservation status of this          threatened species. Environmental Biology of Fishes 87: 55-88.   PDF

Pruitt, J.N., S.E. Riechert, G. Iturralde, M. Vega, B.M. Fitzpatrick, and L. Aviles. 2010. Population differences in          behaviour are explained by shared within-population trait correlations. Journal of Evolutionary Biology

Duncan, S.I., S.E. Riechert, B.M. Fitzpatrick, and J.A. Fordyce. 2010. Relatedness and genetic structure in a socially          polymorphic population of the spider Anelosimus studiosus. Molecular Ecology 19: 810-818.   PDF


Fitzpatrick, B. M., J. A. Fordyce, and S. Gavrilets. 2009. Pattern, process and geographic modes of speciation.          Journal of Evolutionary Biology 22: 2342-2347.   PDF

Fitzpatrick, B. M.. 2009. Power and sample size for nested analysis of molecular variance. Molecular Ecology 18:          3961-3966.   PDF

Todd-Thompson, M., D.L. Miller, P.E. Super, and M.J. Gray. 2009. Chytridiomycosis-associated mortality in Rana          palustris collected in Great Smoky Mountains National Park, Tennessee, USA. Herpetological Review 40:          321-323.   

Fitzpatrick, B.M., J.R. Johnson, D.K. Kump, H.B. Shaffer, J.J. Smith, and S.R. Voss. 2009. Rapid fixation of non-native          alleles revealed by genome-wide SNP analysis of hybrid tiger salamanders. BMC Evolutionary Biology 9: 176.

Ryan, M. E., J. R. Johnson, and B. M. Fitzpatrick. 2009. Invasive hybrid tiger salamander genotypes impact native          amphibians. Proceedings of the National Academy of Sciences, USA. 106: 11166-11171.   PDF

Fitzpatrick, B. M., K. Shook, and R. Izally . 2009. Frequency-dependant selection by wild birds promotes polymorphism          in model salamanders. BMC Ecology 9:12.   PDF

Pasachnik, S. A., B. M. Fitzpatrick, T. J. Near, and A. C. Echternacht. 2009. Gene flow between an endangered          endemic iguana, and its wide spread relative, on the island of Utila, Honduras: When is hybridization a threat?          Conservation Genetics 10: 1247-1254.   PDF

Niemiller, M. L., B. M. Glorioso, C. Nicholas, J. Phillips, J. Rader, E. Reed, K. L. Sykes, J. Todd, G .R. Wyckoff, E. L.          Young, and B. T. Miller. 2009. Notes on the reproduction of the streamside salamander, Ambystoma barbouri,          from Rutherford County, Tennessee. Southeastern Naturalist 8: 37-44.   PDF

Niemiller, M.L., B.T. Miller, and B.M. Fitzpatrick. 2009. Systematics and evolutionary history of subterranean          Gyrinophilus salamanders. Proceedings of the 15th International Congress of Speleology, Kerrville, Texas          15: 242-248.

Niemiller, M.L., B.T. Miller. 2009. A survey of the cave-associated amphibians of the eastern United States with an          emphasis on salamanders. Proceedings of the 15th International Congress of Speleology, Kerrville, Texas          15: 249-256.


Miller, B.T., M. L. Niemiller, and R. G. Reynolds. 2008. Observations on egg-laying behavior and interactions          among attending female Red Salamanders (Pseudotriton ruber) with comments on the use of caves by this          species. Herpetological Conservation and Biology. 3: 203-210.   PDF

Niemiller, M. L. and B. M. Fitzpatrick. 2008. Phylogenetics of the southern cavefish, Typhlichthys subterraneus:          implications for conservation and management. Proceedings of the 2007 National Cave and Karst          Management Symposium. 79-88.   PDF

Fitzpatrick, B. M., J. A. Fordyce, and S. Gavrilets. 2008. What, if anything, is sympatric speciation? Journal of          Evolutionary Biology 21: 1452-1459.   PDF

Fitzpatrick, B. M., J. S. Placyk, M. L. Niemiller, G. S. Casper, and G. M. Burghardt. 2008. Distinctiveness in the          face of gene flow: Hybridization between specialist and generalist gartersnakes. Molecular Ecology 17:          4107-4117.   PDF

Fitzpatrick, B. M. 2008. Hybrid dysfunction: Population genetic and quantitative genetic perspectives. American          Naturalist 171: 491-498.   PDF

Niemiller, M. L., B. M. Fitzpatrick, and B. T. Miller. 2008. Recent divergence with gene flow in Tennessee cave          salamanders (Plethodontidae: Gyrinophilus) inferred from gene genealogies. Molecular Ecology 17: 2258-2275.          PDF

Miller, B. T. and M. L. Niemiller. 2008. Distribution and relative abundance of Tennessee Cave Salamanders          (Gyrinophilous palleucus and Gyrinophilous gulolineatus) with an emphasis on Tennessee populations.          Herpetological Conservation and Biology 3: 1-20.   PDF

Fitzpatrick, B. M. 2008. Dobzhansky-Muller model of hybrid dysfunction supported by poor burst-speed performance in          hybrid tiger salamanders. Journal of Evolutionary Biology. 21: 342-351.   PDF


Bolnick, D. I. and B. M. Fitzpatrick. 2007. Sympatric speciation: models and empirical evidence. Annual Review of          Ecology, Evolution, and Systematics. 38: 459-487.   PDF

Fitzpatrick, B. M. and H. B. Shaffer. 2007. Hybrid vigor between native and introduced salamanders raises new          challenges for conservation. Proceedings of the National Academy of Sciences, USA. 104: 15793-15798.  PDF

McAllister, C. T., C. R. Bursey, M. L. Niemiller, and B. T. Miller. 2007. A noteworthy infection of Clinostomum          complanatum (Digenea: Clinostomidae) in a cave salamander, Eurycea lucifiga (Caudata: Plethodontidae), from          Northcentral Tennessee. Texas Journal of Science 59: 326.  PDF

Reynolds, R. G. and B. M. Fitzpatrick. 2007. Assortative mating in poison-dart frogs based on an ecologically          important trait. Evolution. 61: 2253-2259.  PDF

Niemiller, M. L. and B. T. Miller. 2007. Subterranean reproduction of the Southern Two-lined Salamander (Eurycea          cirrigera) from Short Mountain, Tennessee. Herpetological Conservation and Biology. 2: 106-112.   PDF

Fitzpatrick, B. M. and H. B. Shaffer. 2007. Introduction history and habitat variation explain the landscape genetics of          hybrid tiger salamanders. Ecological Applications. 17: 598-608.   PDF


Fitzpatrick, B. M. and M. Turelli. 2006. The geography of mammalian speciation: mixed signals from phylogenies and          range maps. Evolution. 60: 601-615.   PDF

Niemiller M. L., B. M. Glorioso, C. Nicholas, J. Phillips, J. Rader, E. Reed, K. L. Sykes, J. Todd, G. R. Wyckoff, E. L.          Young, and B. T. Miller. 2006. Status and distribution of the streamside salamander (Ambystoma barbouri) in          middle Tennessee. American Midland Naturalist. 156: 393–399.   PDF

Glorioso B. M., and M. L. Niemiller. 2006. Using deep-water crawfish nets to capture aquatic turtles. Herpetological          Review. 37: 185–187  PDF


Pasachnik, S. and G. R. Ruthig. 2004. Versatility of habitat use in three sympatric species of Plethodontid salamanders.          Journal of Herpetology. 38: 434-437.   PDF

Fitzpatrick, B. M. 2004. Rates of evolution of hybrid inviability in birds and mammals. Evolution. 58: 1865-1870.   PDF

Fitzpatrick, B. M. & H. B. Shaffer. 2004. Environment-dependent admixture dynamics in a tiger salamander hybrid zone.          Evolution. 58: 200-211.  PDF


Riley, S. P. D., H. B. Shaffer, S. R. Voss, and B. M. Fitzpatrick. 2003. Hybridization between a rare, native tiger          salamander (Ambystoma californiense) and its introduced congener. Ecological Applications. 13: 1263-1275.          PDF

Fitzpatrick, B. M., M. F. Benard, J. A. Fordyce. 2003. Morphology and escape performance of tiger salamander larvae          (Ambystoma tigrinum mavortium). Journal of Experimental Zoology. 297A: 147-159.   PDF


Fitzpatrick, B. M. 2002. Molecular correlates of reproductive isolation. Evolution. 56: 191-198.   PDF


Fitzpatrick, B. M. & J. R. Dunk. 1999. Ecogeographic variation in red-tailed hawks. Journal of Raptor Research. 33:          305-312.

2010 All images are the property of R. Graham Reynolds and Matt Niemiller